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Generalized Lotka–Volterra equation : ウィキペディア英語版
Generalized Lotka–Volterra equation
The generalized Lotka–Volterra equations are a set of equations which are more general than either the competitive or predator-prey examples of Lotka–Volterra types.〔
〕 They can be used to model direct competition and trophic relationships between an arbitrary number of species. Their dynamics can be analysed analytically to some extent. This makes them useful as a theoretical tool for modeling food webs. However, they lack features of other ecological models such as predator preference and nonlinear functional responses, and they cannot be used to model mutualism without allowing indefinite population growth.
The Generalised Lotka-Volterra equations model the dynamics of the populations x_1, x_2, \dots of n biological species. Together, these populations can be considered as a vector \mathbf. They are a set of ordinary differential equations given by
:
\frac = x_i f_i(\mathbf),

where the vector \mathbf is given by
:
\mathbf = \mathbf + A\mathbf,

where \mathbf is a vector and A is a matrix known as the community matrix.
==Meaning of parameters==

The generalised Lotka-Volterra equations can represent competition and predation, depending on the values of the parameters, as described below. They are less suitable for describing mutualism.
The values of \mathbf are the intrinsic birth or death rates of the species. A positive value for r_i means that species i is able to reproduce in the absence of any other species (for instance, because it is a plant), whereas a negative value means that its population will decline unless the appropriate other species are present (e.g. a herbivore that cannot survive without plants to eat, or a predator that cannot persist without its prey).
The values of the matrix A represent the relationships between the species. The value of a_ represents the effect that species j has upon species i. The effect is proportional to the populations of both species, as well as to the value of a_. Thus, if both a_ and a_ are negative then the two species are said to be in direct competition with one another, since they each have a direct negative effect on the other's population. If a_ is positive but a_ is negative then species i is considered to be a predator (or parasite) on species j, since i's population grows at j's expense.
Positive values for both a_ and a_ would be considered mutualism. However, this is not often used in practice, because it can make it possible for both species' populations to grow indefinitely.
Indirect negative and positive effects are also possible. For example, if two predators eat the same prey then they compete indirectly, even though they might not have a direct competition term in the community matrix.
The diagonal terms a_ are usually taken to be negative (i.e. species i's population has a negative effect on itself). This self-limitation prevents populations from growing indefinitely.

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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